Discovery Fellows Alumni

Program Alumni Who Have Completed Their PhDs

Daniel Clark, DDS, PhD

Oral and Craniofacial Sciences

Daniel is continuing his training at UCSF as a postdoctoral scholar.

Scott Coyle, PhD


Scott was a 2015 recipient of the prestigious Harold M. Weintraub Graduate Student Award from the Fred Hutchinson Cancer Research Center. He is currently an assistant professor of biochemistry at University of Wisconsin, Madison.

Kelly Crotty, PhD


Kelly is working at the National Cancer Institute as a communications fellow within the Center for Strategic Scientific Initiatives. 

Adolfo Cuesta, PhD

Chemistry and Chemical Biology

A member of UCSF's Medical Scientist Training Program, Adolfo has returned to medical school.

Ryan Dalton, PhD


In 2015, Ryan was awarded a Miller Research Fellowship to do postdoctoral research at UC Berkeley with Dr. Diana Bautista.

Yvanka de Soysa, PhD

Yvanka de Soysa, PhD

Biomedical Sciences

Yvanka is a postdoctoral research fellow at Harvard Medical School/Boston Children's Hospital.

Nima Emami, PhD

Biological and Medical Informatics

Nima is co-founder and CEO of Avail Bio, an SF-based startup focused on creating new products for precision medicine in cancer treatment.

D'Juan Farmer, PhD


D'Juan is currently a postdoc at USC, working with mentor Gage Crump, whose research focuses on craniofacial development. In 2018, he was awarded a prestigious HHMI Hanna H. Gray Fellowship to support his research and training.

Jonathan Gable, PhD


Jonathan is currently an investigator at the Novartis Institute for Tropical Diseases at the Novartis Institutes for Biomedical Research.

John Gagnon, PhD

Biomedical Sciences

John is a scientist at PACT Pharma, where he's conducting research in immunology with a specialization in T cell biology.

Matthew George, PhD

Matthew George, PhD

Developmental and Stem Cell Biology

Matthew is associate director of R&D at Vascugen in Madison, WI.

Allison Girasole, PhD


Allison is doing a postdoc at Harvard Medical School in the lab or Bernardo Sabatini.

Juliet Goldsmith, PhD

Juliet Goldsmith, PhD

Biomedical Sciences

Juliet is a postdoctoral researcher at the Perelman School of Medicine at UPenn with mentor Erika Holzbauer.

Srijib Goswami

Pharmaceutical Science and Pharmacogenomics

Sirj is founder/CEO of Insight RX, a startup focused on leveraging pharmacometrics in a cloud-based software platform to enable precision dosing for healthcare practitioners in real time.

Charles Greenberg, PhD


Charles is currently a health sensing data scientist at Apple, where he works on building automated approaches for interpreting health data from sensors.


Program Alumni Completing their Studies at UCSF

Tara Aitken, Neuroscience     
Francesca Aloisio, Biomedical Sciences    
Varun Bhadkamkar, Tetrad     
Ryan Boileau, Developmental and Stem Cell Biology 
Cassandra Burnett, Biomedical Sciences    
Taylor Cavazos, Biological and Medical Informatics  
Emmalyn Chen, Pharmaceutical Science and Pharmacogenomics  
Jiapei Chen, Biomedical Sciences    
Marcus Chin, Pharmaceutical Sciences and Pharmacogenomics  
Frances Cho, Neuroscience     
Alison Comrie, Neuroscience     
Adam Cotton, Chemistry and Chemical Biology  
Olivia Creasey, Bioengineering     
Shizhong Dai, Chemistry and Chemical Biology  
Sean de la O, Developmental and Stem Cell Biology
Devan Diwanji, Biomedical Sciences    
Susanna Elledge, Chemistry and Chemical Biology  
Ramin Farhad, Oral and Craniofacial Sciences  
Kyle Fowler, Tetrad     
Parinaz Fozouni, Biomedical Sciences    
Nathan Gamarra, Tetrad     
Muryam Gourdet, Tetrad     
Nairi Hartooni, Tetrad     
Clara Herrera, Tetrad     
Joe Hiatt, Biomedical Sciences    
Austin Hsu, Biomedical Sciences    
Kaitlin Hulce, Chemistry and Chemical Biology  
Matthew Jones, Biological and Medical Informatics  
Madeline Keenen, Tetrad     
Nadja Kern, Biophysics     
Eszter Kish, Neuroscience     
Lay Kodama, Neuroscience     
Hardik Kothare, Bioengineering     
Nicole Koutsodendris, Developmental and Stem Cell Biology 
Alexis Krup, Biomedical Sciences    
Kristoffer Leon, Biomedical Sciences    
Lina Leon, Tetrad     
Greyson Lewis, Biophysics     
Anna Lipkin, Neuroscience     
Kyle Lopez, Biophysics     
Benjamin Mansky, Neuroscience     
Christopher Mathy, Bioengineering     
Sarah McMahon, Biomedical Sciences    
Nadia Mohammed Elmassalami Ayad, Bioengineering   
Priya Mohindra, Bioengineering     
Jonathon Muncie, Bioengineering     
Elise Munoz, Tetrad     
Tamas Nagy, Biological and Medical Informatics  
Lila Neahring, Developmental and Stem Cell Biology 
An Nguyen, Oral and Craniofacial Sciences  
Stefan Niekamp, Biophysics     
Carlos Nowotny, Tetrad     
Lindsay Osso, Neuroscience     
Xingjie Pan, Bioengineering     
Ariane Panzer, Biomedical Sciences    
Ramiro Patino, Biomedical Sciences    
Karina Perlaza, Tetrad     
Neha Prasad, Chemistry and Chemical Biology  
Kendra Radtke, Pharmaceutical Sciences and Pharmacogenomics  
Inez Raharjo, Bioengineering     
Capria Rinaldi, Pharmaceutical Sciences and Pharmacogenomics  
Jonathan Schor, Neuroscience     
Charles Schurman, Bioengineering     
Eric Simental, Tetrad     
Taylor Skokan, Tetrad     
Paola Soto-Perez, Tetrad     
Sara Sunshine, Biomedical Sciences    
Serena Tamura, Pharmaceutical Sciences and Pharmacogenomics  
Ngoc-Han Thi Tran, Tetrad    
Ryan Tibble, Chemistry and Chemical Biology  
Kristine Trotta, Tetrad     
Melissa Truong, Developmental and Stem Cell Biology 
Kaitlyn Tsai, Chemistry and Chemical Biology  
Fatima Ugur, Chemistry and Chemical Biology  
Ivana Vasic, Bioengineering     
Sara Vazquez, Tetrad     
Chase Webb, Pharmaceutical Sciences and Pharmacogenomics  
Taia Wu, Chemistry and Chemical Biology  
Xiaowei Yan, Tetrad    

See current Fellows or find information about the Discovery Program.