diverse training in biochemistry and molecular biology, cell biology, developmental biology, and genetics

The Tetrad graduate program prepares students to address fundamental research problems in modern biology, with special emphasis on four core and interrelated areas indicated above. The program as a whole provides interdisciplinary training and opportunities in a congenial community of outstanding research laboratories. When you graduate from the Tetrad program, your PhD degree will be in either biochemistry and molecular biology, cell biology, or genetics, depending on your thesis work. However, all applicants to the program will go through the same admission process, and all students will have the same classes and requirements. Read more about the four research areas within the Tetrad program on the program website.

The Tetrad program fosters a strong sense of community and cross-discipline collaboration through several program activities including an annual retreat held in Lake Tahoe; a weekly seminar program featuring top scientists from around the world; and multiple other activities that provide student/faculty interactions in both formal and informal settings.

Over 120 faculty members are associated with the Tetrad program, representing all of the basic science departments of UCSF; numerous research centers such as the Helen Diller Family Comprehensive Cancer Center, the Cardiovascular Research Institute at UCSF, and the Institute for Neurodegenerative Diseases; as well as numerous clinical departments.

Tetrad is a member of the the Program in Biological Sciences (PIBS) at UCSF.


biochemistry and molecular biology
cell biology

The Tetrad program office is located at the Mission Bay campus, though some classes and labs affiliated with the program are at the Parnassus campus as well. Visit the program website for more information.

The Tetrad program is offered by the UCSF Graduate Division
administered by the UCSF Department of Biochemistry and Biophysics
delivered by faculty members in the UCSF Schools of Dentistry, Medicine, and Pharmacy