Program Alumni Completing their Studies at UCSF
Tara Aitken - Neuroscience
Varun Bhadkamkar - Tetrad
Saumya Bollam - Biomedical Sciences
Emily Bulger - Developmental and Stem Cell Biology
Cassandra Burnett - Biomedical Sciences
Jiapei Chen - Biomedical Sciences
Alison Comrie - Neuroscience
Jessica Cook - Oral and Craniofacial Sciences
Katherine Crawford - Bioengineering
Olivia Creasey - Bioengineering
Sean de la O - Developmental and Stem Cell Biology
Ramin Farhad - Oral and Craniofacial Sciences
Haley Gause - Tetrad
Mollie Eva Hansen - Bioengineering
Scott Harris - Neuroscience
Evelyn Hernandez - Tetrad
Clara Herrera - Tetrad
Matthew Johnson - Biological and Medical Informatics
Eszter Kish - Neuroscience
Jose Liboy - Tetrad
Harold Marin - Tetrad
Christopher Mathy - Bioengineering
Michael Mobaraki - Developmental and Stem Cell Biology
Nadia Mohammed Elmassalami Ayad - Bioengineering
Priya Mohindra - Bioengineering
Elise Munoz - Tetrad
An Nguyen - Oral and Craniofacial Sciences
Carlos Nowotny - Tetrad
Ramiro Patino - Biomedical Sciences
Gokul Ramadoss - Biomedical Sciences
Capria Rinaldi - Pharmaceutical Sciences and Pharmacogenomics
Eric Simental - Tetrad
Christina Stephens - Biophysics
Chase Webb - Pharmaceutical Sciences and Pharmacogenomics
Gina Williams - Neuroscience
Taia Sean Wu - Chemistry and Chemical Biology
Graduates Who Have Completed Their PhDs
Shaun Abrams, PhD in Oral and Craniofacial Sciences (2017)
Osama Ahmed, PhD in Neuroscience (2017)
Ibraheem Ali, PhD in Biomedical Sciences (2018)
Francesca Aloisio, PhD in Biomedical Sciences (2021)
Brittany Anderton, PhD in Biomedical Sciences (2015)
Benjamin Barad, PhD in Biophysics (2019)
Kyle Barlow, PhD in Biological and Medical Informatics (2017)
Ryan Boileau, PhD in Developmental and Stem Cell Biology (2023)
Christine Bowman, PhD in Pharmaceutical Sciences and Pharmacogenomics (2019)
Deanna Brackman, PhD in Pharmaceutical Sciences and Pharmacogenomics (2018)
Kenneth Burke, PhD in Neuroscience (2018)
Roman Camarda, PhD in Biomedical Sciences (2018)
Andrea Cantu, PhD in Biomedical Sciences (2016)
Jessie Carr, PhD in Pharmaceutical Sciences and Pharmacogenomics (2017)
Taylor Cavazos, PhD in Biological and Medical Informatics (2022)
Alec Cerchiari, PhD in Bioengineering (2014)
Amy Chang, PhD in Biophysics (2017)
Emmalyn Chen, PhD in Pharmaceutical Sciences and Pharmacogenomics (2020)
Yiran Chen, PhD in Bioengineering (2018)
Gauree Chendke, PhD in Bioengineering (2023)
Marcus Chin, PhD in Pharmaceutical Sciences and Pharmacogenomics (2021)
Frances Cho, PhD in Neuroscience (2022)
Abrar Choudhury, PhD in Biomedical Sciences (2021)
Daniel Clark, PhD in Oral and Craniofacial Sciences (2019)
Emily Connelly, PhD in Pharmaceutical Sciences and Pharmacogenomics (2023)
Adam Cotton, PhD in Chemistry and Chemical Biology (2021)
Scott Coyle, PhD in Tetrad (2015)
Kelly Crotty, PhD in Tetrad (2019)
Adolfo Cuesta, PhD in Chemistry and Chemical Biology (2019)
Shizhong Dai, PhD in Chemistry and Chemical Biology (2022)
Ryan Dalton, PhD in Neuroscience (2015)
Yvanka de Soysa, PhD in Biomedical Sciences (2019)
Devan Diwanji, PhD in Biomedical Sciences (2021)
Adam Elkhaled, PhD in Bioengineering (2016)
Susanna Elledge, PhD in Chemistry and Chemical Biology (2022)
Nima Emami, PhD in Biological and Medical Informatics (2018)
D'Juan Farmer, PhD in Tetrad (2017)
Kyle Fowler, PhD in Tetrad (2022)
Parinaz Fozouni, PhD in Biomedical Sciences (2020)
Jonathan Gable, PhD in Biophysics (2015)
John Gagnon, PhD in Biomedical Sciences (2018)
Nathan Gamarra, PhD in Tetrad (2020)
Daniel Gentile, PhD in Chemistry and Chemical Biology (2017)
Matthew George, PhD in Developmental and Stem Cell Biology (2018)
Allison Girasole, PhD in Neuroscience (2019)
Juliet Goldsmith, PhD in Biomedical Sciences (2018)
Srijib Goswami, PhD in Pharmaceutical Sciences and Pharmacogenomics (2015)
Muryam Gourdet, PhD in Tetrad (2022)
Charles Greenberg, PhD in Biophysics (2016)
Jennifer Guidera, PhD in Bioengineering (2023)
Nairi Hartooni, PhD in Tetrad (2021)
Benjamin Heineike, PhD in Biological and Medical Informatics (2019)
Joseph Hiatt, PhD in Biomedical Sciences (2020)
Daniel Holohan, PhD in Biomedical Sciences (2019)
Christina Homer, PhD in Biophysics (2016)
Maximilian Horlbeck, PhD in Biophysics (2017)
Austin Hsu, PhD in Biomedical Sciences (2022)
Christina Hueschen, PhD in Biomedical Sciences (2018)
Kaitlin Hulce, PhD in Chemistry and Chemical Biology (2021)
Amy Jacobson, PhD in Chemistry and Chemical Biology (2017)
Arman Jahangiri, PhD in Biomedical Sciences (2018)
Llewellyn "Trey" Jalbert, PhD in Bioengineering (2015)
Ruiji Jiang, PhD in Biomedical Sciences (2019)
Isabel Johnson, PhD in Tetrad (2020)
Emily Jones, PhD in Biomedical Sciences (2019)
Kyle Jones, PhD in Oral and Craniofacial Sciences (2017)
Matthew Jones, PhD in Biological and Medical Informatics (2022)
Ariel Kauss, PhD in Biomedical Sciences (2020)
Madeline Keenen, PhD in Tetrad (2020)
Dyana Kenanova, PhD in Chemistry and Chemical Biology (2023)
Kathleen Keough, PhD in Pharmaceutical Sciences and Pharmacogenomics (2020)
Nadja Kern, PhD in Biophysics (2021)
Natalia Khuri, PhD in Biophysics (2014)
Jacob Kimmel, PhD in Developmental and Stem Cell Biology (2018)
Lay Kodama, PhD in Neuroscience (2020)
Bertram Koelsch, PhD in Bioengineering (2014)
Natalie Korn, PhD in Bioengineering (2019)
Kamena Kostova, PhD in Biomedical Sciences (2018)
Hardik Kothare, PhD in Bioengineering (2020)
Nicole Koutsodendris, PhD in Developmental and Stem Cell Biology (2022)
Laurens Kraal, PhD in Biological and Medical Informatics (2015)
Alexis Krup, PhD in Biomedical Sciences (2022)
Jonathan Kuhn, PhD in Tetrad (2019)
Jennifer Kung, PhD in Chemistry and Chemical Biology (2018)
Taylor LaFlam, PhD in Biomedical Sciences (2015)
Alexis Lainoff, PhD in Oral and Craniofacial Sciences (2019)
Samira Khakpour Lawton, PhD in Biomedical Sciences (2017)
Alison Leaf, PhD in Tetrad (2015)
Andrew Lechner, PhD in Biomedical Sciences (2017)
David Leib, PhD in Neuroscience (2018)
Kristoffer Leon, PhD in Biomedical Sciences (2020)
Lina Leon, PhD in Tetrad (2021)
Greyson Lewis, PhD in Biophysics (2022)
Ashley Libby, PhD in Developmental and Stem Cell Biology (2019)
John Lin King, PhD in Neuroscience (2020)
Robert Lindquist, PhD in Neuroscience (2015)
Anna Lipkin, PhD in Neuroscience (2023)
Harrison Liu, PhD in Bioengineering (2016)
Jennifer Liu, PhD in Biomedical Sciences (2020)
Alexandra Long, PhD in Tetrad (2020)
Jocelyne Lopez, PhD in Tetrad (2023)
Kyle Lopez, PhD in Biophysics (2022)
Andrew Lyon, PhD in Tetrad (2017)
Andrew Mancini, PhD in Biomedical Sciences (2018)
Benjamin Mansky, PhD in Neuroscience (2020)
Florie Mar, PhD in Biomedical Sciences (2015)
Timothy Marsh, PhD in Biomedical Sciences (2019)
Elizabeth McCarthy, PhD in Biological and Medical Informatics (2022)
Justin Mclaurin, PhD in Tetrad (2018)
Sarah McMahon, PhD in Biomedical Sciences (2021)
Karuna Meda, PhD in Neuroscience (2017)
Fernando Meza Gutierrez, PhD in Tetrad (2018)
Eugene Milshteyn, PhD in Bioengineering (2017)
Yekaterina Miroshnikova, PhD in Bioengineering (2015)
Christine Moore Sheridan, PhD in Biomedical Sciences (2018)
Marco Mravic, PhD in Biophysics (2020)
Jonathon Muncie, PhD in Bioengineering (2020)
Tamas Nagy, PhD in Biological and Medical Informatics (2023)
Ramsey Najm, PhD in Developmental and Stem Cell Biology (2019)
Lila Neahring, PhD in Developmental and Stem Cell Biology (2023)
Stefan Niekamp, PhD in Biophysics (2020)
Isabel Nocedal, PhD in Tetrad (2016)
Crystal Nyitray, PhD in Chemistry and Chemical Biology (2015)
Nicole Olson, PhD in Chemistry and Chemical Biology (2016)
Lindsay Osso, PhD in Neuroscience (2021)
Beatriz Osuna, PhD in Tetrad (2019)
Xingjie Pan, PhD in Bioengineering (2020)
Katarina Pance, PhD in Chemistry and Chemical Biology (2021)
Ariane Panzer, PhD in Biomedical Sciences (2021)
David Paquette, PhD in Biophysics (2017)
Danny Park, PhD in Pharmaceutical Sciences and Pharmacogenomics (2017)
Kartik Pattabiraman, PhD in Neuroscience (2014)
Karina Perlaza, PhD in Tetrad (2021)
Veronica Pessino, PhD in Biophysics (2017)
Brenden Petersen, PhD in Bioengineering (2016)
Cameron Pitt, PhD in Biomedical Sciences (2015)
Neha Prasad, PhD in Chemistry and Chemical Biology (2022)
Kendra Radtke, PhD in Pharmaceutical Sciences and Pharmacogenomics (2021)
Inez Raharjo, PhD in Bioengineering (2020)
Hanna Retallack, PhD in Biomedical Sciences (2020)
T. Justin Rettenmaier, PhD in Chemistry and Chemical Biology (2015)
Matthew Rubashkin, PhD in Bioengineering (2016)
Karen Ruiz, PhD in Tetrad (2019)
Florentine Rutaganira, PhD in Chemistry and Chemical Biology (2016)
Dan Santos, PhD in Tetrad (2019)
Erica Schlesinger, PhD in Bioengineering (2015)
Jonathan Schor, PhD in Neuroscience (2020)
Courtney Schroeder, PhD in Tetrad (2016)
Charles Schurman, PhD in Bioengineering (2022)
Hong Shang, PhD in Bioengineering (2016)
Meera Shenoy, PhD in Biomedical Sciences (2017)
Taylor Skokan, PhD in Tetrad (2022)
Geoffrey Smith, PhD in Chemistry and Chemical Biology (2017)
Paola Soto-Perez, PhD in Tetrad (2021)
Perry Spratt, PhD in Neuroscience (2020)
Hanna Starobinets, PhD in Biomedical Sciences (2016)
Caitlin Stoddard, PhD in Tetrad (2018)
Meena Subramaniam, PhD in Biological and Medical Informatics (2019)
Sara Sunshine, PhD in Biomedical Sciences (2022)
Christine Leon Swisher, PhD in Bioengineering (2014)
Serena Tamura, PhD in Pharmaceutical Sciences and Pharmacogenomics (2021)
Fei Tan, PhD in Bioengineering (2023)
Shuyu Tang, PhD in Bioengineering (2019)
Isabelle Taylor, PhD in Chemistry and Chemical Biology (2018)
Christina Theodoris, PhD in Developmental and Stem Cell Biology (2015)
Ryan Tibble, PhD in Chemistry and Chemical Biology (2021)
May Tran, PhD in Neuroscience (2017)
Ngoc-Han Thi Tran, PhD in Tetrad (2020)
Kristine Trotta, PhD in Tetrad (2023)
Melissa Truong, PhD in Developmental and Stem Cell Biology (2021)
Mary Kaitlyn Tsai, PhD in Chemistry and Chemical Biology (2021)
Nicole Tsai, PhD in Biomedical Sciences (2022)
Fatima Ugur, PhD in Chemistry and Chemical Biology (2022)
Lawrence Uricchio, PhD in Biological and Medical Informatics (2014)
Katherine Varandas, PhD in Tetrad (2016)
Ivana Vasic, PhD in Bioengineering (2022)
Sara Vazquez, PhD in Tetrad (2021)
Tess Veuthey, PhD in Neuroscience (2019)
Ivan Vujkovic-Cvijin, PhD in Biomedical Sciences (2015)
Chenyu Wang, PhD in Neuroscience (2023)
Stephanie Wankowicz, PhD in Biophysics (2023)
Racheli Wercberger, PhD in Neuroscience (2019)
Peter Westcott, PhD in Pharmaceutical Sciences and Pharmacogenomics (2015)
Jasper Williams, PhD in Pharmaceutical Sciences and Pharmacogenomics (2019)
Jessica Witchley, PhD in Tetrad (2018)
Alex Wolff, PhD in Biophysics (2020)
Sarah Wong, PhD in Oral and Craniofacial Sciences (2018)
Xiaowei Yan, PhD in Tetrad (2021)
Julia Ye, PhD in Biomedical Sciences (2017)
Xucheng Zhu, PhD in Bioengineering (2020)
Christopher Zimmerman, PhD in Neuroscience (2019)
Meghan Zubradt, PhD in Biomedical Sciences (2016)
See current Fellows or find information about the Discovery Program.
