Program Alumni Completing their Studies at UCSF
Hersh Bhargava, Biophysics
Lauren Cech, Pharmaceutical Sciences and Pharmacogenomics
Nathan Cho, Tetrad
Cambria Chou-Freed, Biomedical Sciences
Jessica Cook - Oral and Craniofacial Sciences
Rebecca Fang, Neuroscience
Mollie Eva Hansen - Bioengineering
Jordan Kleinman, Chemistry and Chemical Biology
Monique Lillis, Neuroscience
Carlos Nowotny - Tetrad
Ali Palla, Tetrad
Tiange (Tony) Qu, Oral and Craniofacial Sciences
Elizabeth Sisko, Chemistry and Chemical Biology
Cameron Wilhite, Neuroscience
Muziyue Wu, Biophysics
Jenny Zhang, Tetrad
Graduates Who Have Completed Their PhDs
A
Shaun Abrams, PhD in Oral and Craniofacial Sciences (2017)
Osama Ahmed, PhD in Neuroscience (2017)
Tara Aitken, PhD in Neuroscience (2023)
Ibraheem Ali, PhD in Biomedical Sciences (2018)
Francesca Aloisio, PhD in Biomedical Sciences (2021)
Brittany Anderton, PhD in Biomedical Sciences (2015)
Francisco Aparicio, PhD in Neuroscience (2024)
B
Benjamin Barad, PhD in Biophysics (2019)
Kyle Barlow, PhD in Biological and Medical Informatics (2017)
Varun Bhadkamkar, PhD in Tetrad (2023)
Ryan Boileau, PhD in Developmental and Stem Cell Biology (2023)
Saumya Bollam, PhD in Biomedical Sciences (2024)
Elizabeth Bond, PhD in Tetrad (2024)
Christine Bowman, PhD in Pharmaceutical Sciences and Pharmacogenomics (2019)
Deanna Brackman, PhD in Pharmaceutical Sciences and Pharmacogenomics (2018)
Emily Bulger, PhD in Developmental and Stem Cell Biology (2024)
Kenneth Burke, PhD in Neuroscience (2018)
Cassandra Burnett, PhD in Biomedical Sciences (2023)
C
Roman Camarda, PhD in Biomedical Sciences (2018)
Andrea Cantu, PhD in Biomedical Sciences (2016)
Jessie Carr, PhD in Pharmaceutical Sciences and Pharmacogenomics (2017)
Taylor Cavazos, PhD in Biological and Medical Informatics (2022)
Alec Cerchiari, PhD in Bioengineering (2014)
Amy Chang, PhD in Biophysics (2017)
Emmalyn Chen, PhD in Pharmaceutical Sciences and Pharmacogenomics (2020)
Jiapei Chen, PhD in Biomedical Sciences (2023)
Gauree Chendke, PhD Bioengineering (2023)
Yiran Chen, PhD in Bioengineering (2018)
Gauree Chendke, PhD in Bioengineering (2023)
Marcus Chin, PhD in Pharmaceutical Sciences and Pharmacogenomics (2021)
Frances Cho, PhD in Neuroscience (2022)
Abrar Choudhury, PhD in Biomedical Sciences (2021)
Daniel Clark, PhD in Oral and Craniofacial Sciences (2019)
Mitchel Cole, PhD in Biomedical Sciences (2025)
Alison Comrie, PhD in Neuroscience (2024)
Emily Connelly, PhD in Pharmaceutical Sciences and Pharmacogenomics (2023)
Adam Cotton, PhD in Chemistry and Chemical Biology (2021)
Scott Coyle, PhD in Tetrad (2015)
Katherine Crawford, PhD in Bioengineering (2024)
Olivia Creasey, PhD in Bioengineering (2024)
Kelly Crotty, PhD in Tetrad (2019)
Adolfo Cuesta, PhD in Chemistry and Chemical Biology (2019)
Zachary Cutts, PhD in Biological and Medical Informatics (2024)
D
Shizhong Dai, PhD in Chemistry and Chemical Biology (2022)
Ryan Dalton, PhD in Neuroscience (2015)
Sean de la O, PhD in Developmental and Stem Cell Biology (2023)
Yvanka de Soysa, PhD in Biomedical Sciences (2019)
Devan Diwanji, PhD in Biomedical Sciences (2021)
E
Adam Elkhaled, PhD in Bioengineering (2016)
Susanna Elledge, PhD in Chemistry and Chemical Biology (2022)
Nima Emami, PhD in Biological and Medical Informatics (2018)
F
Ramin Farhad, PhD in Oral and Craniofacial Sciences (2025)
D'Juan Farmer, PhD in Tetrad (2017)
Kyle Fowler, PhD in Tetrad (2022)
Parinaz Fozouni, PhD in Biomedical Sciences (2020)
G
Jonathan Gable, PhD in Biophysics (2015)
Margaret Gadek, PhD in Biomedical Sciences (2025)
John Gagnon, PhD in Biomedical Sciences (2018)
Nathan Gamarra, PhD in Tetrad (2020)
Haley Gause, PhD in Tetrad (2024)
Eliza Gaylord, PhD in Developmental and Stem Cell Biology (2025)
Daniel Gentile, PhD in Chemistry and Chemical Biology (2017)
Matthew George, PhD in Developmental and Stem Cell Biology (2018)
Allison Girasole, PhD in Neuroscience (2019)
Juliet Goldsmith, PhD in Biomedical Sciences (2018)
Srijib Goswami, PhD in Pharmaceutical Sciences and Pharmacogenomics (2015)
Muryam Gourdet, PhD in Tetrad (2022)
Charles Greenberg, PhD in Biophysics (2016)
Jennifer Guidera, PhD in Bioengineering (2023)
Amy Guo, PhD in Bioengineering (2024)
H
Karissa Hansen, PhD in Developmental and Stem Cell Biology (2025)
Scott Harris, PhD in Neuroscience (2024)
Nairi Hartooni, PhD in Tetrad (2021)
Benjamin Heineike, PhD in Biological and Medical Informatics (2019)
Evelyn Hernandez, PhD in Tetrad (2023)
Clara Herrera, PhD in Tetrad (2024)
Joseph Hiatt, PhD in Biomedical Sciences (2020)
Daniel Holohan, PhD in Biomedical Sciences (2019)
Christina Homer, PhD in Biophysics (2016)
Maximilian Horlbeck, PhD in Biophysics (2017)
Austin Hsu, PhD in Biomedical Sciences (2022)
Christina Hueschen, PhD in Biomedical Sciences (2018)
Erin Huiting, PhD in Biomedical Sciences (2024)
Kaitlin Hulce, PhD in Chemistry and Chemical Biology (2021)
J
Amy Jacobson, PhD in Chemistry and Chemical Biology (2017)
Arman Jahangiri, PhD in Biomedical Sciences (2018)
Llewellyn "Trey" Jalbert, PhD in Bioengineering (2015)
Owen Jiang, PhD in Biomedical Sciences (2024)
Ruiji Jiang, PhD in Biomedical Sciences (2019)
Isabel Johnson, PhD in Tetrad (2020)
Matthew Johnson, PhD in Biological and Medical Informatics (2023)
Ellery Jones, PhD in Neuroscience (2025)
Emily Jones, PhD in Biomedical Sciences (2019)
Kyle Jones, PhD in Oral and Craniofacial Sciences (2017)
Matthew Jones, PhD in Biological and Medical Informatics (2022)
Ziad Jowhar, PhD in Biomedical Sciences (2024)
K
Ariel Kauss, PhD in Biomedical Sciences (2020)
Madeline Keenen, PhD in Tetrad (2020)
Dyana Kenanova, PhD in Chemistry and Chemical Biology (2023)
Kathleen Keough, PhD in Pharmaceutical Sciences and Pharmacogenomics (2020)
Nadja Kern, PhD in Biophysics (2021)
Natalia Khuri, PhD in Biophysics (2014)
Jacob Kimmel, PhD in Developmental and Stem Cell Biology (2018)
Ziggy Kish, PhD in Neuroscience (2025)
Lay Kodama, PhD in Neuroscience (2020)
Bertram Koelsch, PhD in Bioengineering (2014)
Natalie Korn, PhD in Bioengineering (2019)
Kamena Kostova, PhD in Biomedical Sciences (2018)
Hardik Kothare, PhD in Bioengineering (2020)
Nicole Koutsodendris, PhD in Developmental and Stem Cell Biology (2022)
Laurens Kraal, PhD in Biological and Medical Informatics (2015)
Alexis Krup, PhD in Biomedical Sciences (2022)
Jonathan Kuhn, PhD in Tetrad (2019)
Jennifer Kung, PhD in Chemistry and Chemical Biology (2018)
L
Dianne Laboy-Cintron, PhD in Biomedical Sciences (2024)
Taylor LaFlam, PhD in Biomedical Sciences (2015)
Alexis Lainoff, PhD in Oral and Craniofacial Sciences (2019)
Samira Khakpour Lawton, PhD in Biomedical Sciences (2017)
Alison Leaf, PhD in Tetrad (2015)
Andrew Lechner, PhD in Biomedical Sciences (2017)
David Leib, PhD in Neuroscience (2018)
Fredrick Leon, PhD in Tetrad (2025)
Kristoffer Leon, PhD in Biomedical Sciences (2020)
Lina Leon, PhD in Tetrad (2021)
Greyson Lewis, PhD in Biophysics (2022)
Yaqiao Li, PhD in Pharmaceutical Sciences and Pharmacogenomics (2024)
Ashley Libby, PhD in Developmental and Stem Cell Biology (2019)
Jose Liboy, PhD in Tetrad (2024)
John Lin King, PhD in Neuroscience (2020)
Robert Lindquist, PhD in Neuroscience (2015)
Anna Lipkin, PhD in Neuroscience (2023)
Harrison Liu, PhD in Bioengineering (2016)
Jennifer Liu, PhD in Biomedical Sciences (2020)
Alexandra Long, PhD in Tetrad (2020)
Jocelyne Lopez, PhD in Tetrad (2023)
Kyle Lopez, PhD in Biophysics (2022)
Andrew Lyon, PhD in Tetrad (2017)
M
Andrew Mancini, PhD in Biomedical Sciences (2018)
Benjamin Mansky, PhD in Neuroscience (2020)
Florie Mar, PhD in Biomedical Sciences (2015)
Harold Marin, PhD in Tetrad (2024)
Timothy Marsh, PhD in Biomedical Sciences (2019)
Christopher Mathy, PhD in Bioengineering (2022)
Elizabeth McCarthy, PhD in Biological and Medical Informatics (2022)
Justin Mclaurin, PhD in Tetrad (2018)
Sarah McMahon, PhD in Biomedical Sciences (2021)
Karuna Meda, PhD in Neuroscience (2017)
Brenda Melano-Gonzales, PhD in Pharmaceutical Sciences and Pharmacogenomics (2025)
Shreya Menon, PhD in Biological and Medical Informatics (2025)
Fernando Meza Gutierrez, PhD in Tetrad (2018)
Eugene Milshteyn, PhD in Bioengineering (2017)
Yekaterina Miroshnikova, PhD in Bioengineering (2015)
Michael Mobaraki, PhD in Developmental and Stem Cell Biology (2023)
Nadia Mohammed Elmassalami Ayad, PhD in Bioengineering (2023)
Priya Mohindra, PhD in Bioengineering (2022)
Christine Moore Sheridan, PhD in Biomedical Sciences (2018)
Marco Mravic, PhD in Biophysics (2020)
Duncan Muir, PhD in Biophysics (2025)
Jonathon Muncie, PhD in Bioengineering (2020)
Elise Munoz, PhD in Tetrad (2024)
N
Tamas Nagy, PhD in Biological and Medical Informatics (2023)
Ramsey Najm, PhD in Developmental and Stem Cell Biology (2019)
Lila Neahring, PhD in Developmental and Stem Cell Biology (2023)
An Nguyen, PhD in Oral and Craniofacial Sciences (2023)
Stefan Niekamp, PhD in Biophysics (2020)
Isabel Nocedal, PhD in Tetrad (2016)
Crystal Nyitray, PhD in Chemistry and Chemical Biology (2015)
O
Nicole Olson, PhD in Chemistry and Chemical Biology (2016)
Lindsay Osso, PhD in Neuroscience (2021)
Beatriz Osuna, PhD in Tetrad (2019)
P
Xingjie Pan, PhD in Bioengineering (2020)
Katarina Pance, PhD in Chemistry and Chemical Biology (2021)
Ariane Panzer, PhD in Biomedical Sciences (2021)
David Paquette, PhD in Biophysics (2017)
Danny Park, PhD in Pharmaceutical Sciences and Pharmacogenomics (2017)
Ramiro Patino, PhD in Biomedical Sciences (2024)
Kartik Pattabiraman, PhD in Neuroscience (2014)
Karina Perlaza, PhD in Tetrad (2021)
Veronica Pessino, PhD in Biophysics (2017)
Brenden Petersen, PhD in Bioengineering (2016)
Cameron Pitt, PhD in Biomedical Sciences (2015)
Neha Prasad, PhD in Chemistry and Chemical Biology (2022)
R
Kendra Radtke, PhD in Pharmaceutical Sciences and Pharmacogenomics (2021)
Inez Raharjo, PhD in Bioengineering (2020)
Gokul Ramadoss, PhD in Biomedical Sciences (2024)
Hanna Retallack, PhD in Biomedical Sciences (2020)
T. Justin Rettenmaier, PhD in Chemistry and Chemical Biology (2015)
Capria Rinaldi, PhD in Pharmaceutical Sciences and Pharmacogenomics (2023)
Jacquelyn Roger, PhD in Biological and Medical Informatics (2025)
Matthew Rubashkin, PhD in Bioengineering (2016)
Karen Ruiz, PhD in Tetrad (2019)
Florentine Rutaganira, PhD in Chemistry and Chemical Biology (2016)
S
Sule Sahin, PhD in Bioengineering (2024)
Dan Santos, PhD in Tetrad (2019)
Erica Schlesinger, PhD in Bioengineering (2015)
Jonathan Schor, PhD in Neuroscience (2020)
Courtney Schroeder, PhD in Tetrad (2016)
Charles Schurman, PhD in Bioengineering (2022)
Hong Shang, PhD in Bioengineering (2016)
Wenqi Shen, PhD in Chemistry and Chemical Biology (2025)
Meera Shenoy, PhD in Biomedical Sciences (2017)
Eric Simental, PhD in Tetrad (2024)
Taylor Skokan, PhD in Tetrad (2022)
Geoffrey Smith, PhD in Chemistry and Chemical Biology (2017)
Paola Soto-Perez, PhD in Tetrad (2021)
Perry Spratt, PhD in Neuroscience (2020)
Hanna Starobinets, PhD in Biomedical Sciences (2016)
Christina Stephens, PhD in Biophysics (2024)
Caitlin Stoddard, PhD in Tetrad (2018)
Meena Subramaniam, PhD in Biological and Medical Informatics (2019)
Sara Sunshine, PhD in Biomedical Sciences (2022)
Christine Leon Swisher, PhD in Bioengineering (2014)
T
Serena Tamura, PhD in Pharmaceutical Sciences and Pharmacogenomics (2021)
Fei Tan, PhD in Bioengineering (2023)
Alice Tang, PhD in Bioengineering (2023)
Shuyu Tang, PhD in Bioengineering (2019)
Isabelle Taylor, PhD in Chemistry and Chemical Biology (2018)
Olivia Teter, PhD in Bioengineering (2025)
Christina Theodoris, PhD in Developmental and Stem Cell Biology (2015)
Ryan Tibble, PhD in Chemistry and Chemical Biology (2021)
May Tran, PhD in Neuroscience (2017)
Ngoc-Han Thi Tran, PhD in Tetrad (2020)
Kristine Trotta, PhD in Tetrad (2023)
Melissa Truong, PhD in Developmental and Stem Cell Biology (2021)
Mary Kaitlyn Tsai, PhD in Chemistry and Chemical Biology (2021)
Nicole Tsai, PhD in Biomedical Sciences (2022)
U
Fatima Ugur, PhD in Chemistry and Chemical Biology (2022)
Lawrence Uricchio, PhD in Biological and Medical Informatics (2014)
V
Katherine Varandas, PhD in Tetrad (2016)
Ivana Vasic, PhD in Bioengineering (2022)
Sara Vazquez, PhD in Tetrad (2021)
Tess Veuthey, PhD in Neuroscience (2019)
Regan Volk, PhD in Chemistry and Chemical Biology (2024)
Ivan Vujkovic-Cvijin, PhD in Biomedical Sciences (2015)
W
Hailey Wallace, PhD in Biophysics (2023)
Chenyu Wang, PhD in Neuroscience (2023)
Stephanie Wankowicz, PhD in Biophysics (2023)
Chase Webb, PhD in Pharmaceutical Sciences and Pharmacogenomics (2023)
Racheli Wercberger, PhD in Neuroscience (2019)
Peter Westcott, PhD in Pharmaceutical Sciences and Pharmacogenomics (2015)
Gina Williams, PhD in Neuroscience (2025)
Jasper Williams, PhD in Pharmaceutical Sciences and Pharmacogenomics (2019)
Jessica Witchley, PhD in Tetrad (2018)
Alex Wolff, PhD in Biophysics (2020)
Sarah Wong, PhD in Oral and Craniofacial Sciences (2018)
Taia Wu, PhD in Chemistry and Chemical Biology (2023)
Y
Xiaowei Yan, PhD in Tetrad (2021)
Julia Ye, PhD in Biomedical Sciences (2017)
Z
Xucheng Zhu, PhD in Bioengineering (2020)
Christopher Zimmerman, PhD in Neuroscience (2019)
Meghan Zubradt, PhD in Biomedical Sciences (2016)
See current Fellows or find information about the Discovery Program.